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To evaluate the nature of the ASPL-VCP interaction, we used a combination of fluorescence-detected sedimentation velocity analytical ultracentrifugation (FDS-SV-AUC) and interference SV-AUC experiments with a 1:84 ratio of ASPL to VCP monomers [0.2 M Alexa Fluor 488 (AF488)ASPL to 16.8 M VCP] to approximate physiological conditions