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Plasmid identity was verified using Sanger sequencing, and the most efficient sgRNA (as determined by anti-GCDH immunoblotting) was used in subsequent experiments (sgRNA sequences are listed in Supplementary Table 1)

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The expression of BOLA3 is reduced due to HIF-2-dependent transcriptional inhibition, and its deletion disrupts the integrity of iron-sulfur (Fe-S) clusters, affects the function of two-oxoacid dehydrogenases containing lipoic acid, and regulates glycolysis and mitochondrial respiration